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Hoppy, as he was known to all, studied natural sciences at Cambridge, followed by clinical training at the London Hospital. Inspired by biochemistry in Cambridge he moved to lecturing and research immediately after completing house jobs. He developed a passionate interest in human genetic variation in relation to both healthy individuals and human disease. From the London Hospital he moved to King’s College and then University College London (UCL). He directed the Medical Research Council (MRC) Human Biochemical Genetics Unit at UCL from 1976 to 2000.